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Try disabling Instant ON on both and see what happens...
I get OPEN NAT on both my Xbox One's without having to disable INSTANT ON
HOW? Please BiggShooter do tell! I'd love to use my instant on but I get strict on my second console as well.
How do you have your XB1s connected to the 5500?
I've seen issues with using DAPs, what happens if you wire direct if possible the 2nd XB1. Regarding DAPs, they can have a non-transparent IP address that appears as a 2ndary IP address. I notice this with my 1533, 865L and 868L in bridge mode. Even though I manually set a static IP address on each, the main host router assigns a different one that I see under the Dynamic Clients list. I presume that this may or may not have some impact on NAT.